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cysteine sulfinic acid organic compound amino functional group enantiomer cysteine metabolism post-translationally acid reductase cystathionine gamma-lyase enzyme lyase dioxygenase taurine 3d metabolite biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous contractor human metabolome database hmdb tautomer isomer stereoisomer orbital structure activity relationship sar racemate substance matter material component