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cofactor f430 enzyme methyl coenzyme reductase mcr coenzyme cofactor methanogenesis methanogenic archaea methanobacterium porphyrin macrocyclic nickel uroporphyrinogen iii biosynthesis cobalamin non-protein catalyst chemical reaction biochemical inorganic ion cof430 organic metabolite metabolism cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous contractor microorganism strain 3d molecule molecular formula