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coenzyme methyl-transfer archaeal methanogens 2-mercaptoethylsulfonate 2-mercaptoethanesulfonate anion h-s-com 2-sulfanylethane-1-sulfonate methanogenesis methane prosthetic group xanthobacter autotrophicus cofactor f430 metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor cell biology biochemistry biomolecule bioactive microorganism strain molecule molecular formula skeletal structural structure geometry 3d chemical