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chlorophyllide oxygenic photosynthesis chlorophyll synthase universal eukaryotes bacteriochlorophyll plants algae cyanobacteria photosynthetic organisms tetrapyrrole chlorophyl chlorophyll pigment chloroplasts algae energy light electromagnetic spectrum wavelengths photosystems thylakoid resonance energy transfer reaction center absorb photosystem ii electron transport chain light nadph porphyrin chlorin magnesium 3d phytyl biosynthesis chlorophyllide photosensitizer cycle molecule molecular formula skeletal structural structure