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protochlorophyllide monovinyl intermediate biosynthesis chlorophyll phytol reduced pyrrole fluorescent angiosperm light-dependent chlorotic gymnosperms algae photosynthetic bacteria light-independent enzyme green darkness chlorophyllide protochlorophyllide reductase flu biosynthesis regulatory protein herbal phytochemical phytotherapy plant herb natural herbalism pharmacognosy pharmacopoeia metabolite metabolism mechanism pathway path biosynthetic biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous contractor 3d