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astaxanthin keto-carotenoid terpenes zeaxanthin canthaxanthin pigment microalgae haematococcus pluvialis yeast fungus xanthophyllomyces dendrorhous dietary supplement red-coloured plankton krill biosynthesis seafood metabolic engineering salmonids xanthophylls molecule molecular formula skeletal structural structure geometry ball stick metabolite metabolism cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous 3d