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5-diphosphomevalonic diphosphomevalonic acid intermediate mevalonate pathway isoprenoid metabolic eukaryotes archaea acetyl coa acetoacetyl amino smpdb hydroxy dihydrolanosterol acetyltransferase synthase reductase methylzymosterol methylthiobutanoic 3d rendering chemical cholesterol steroid biochemistry biology biomolecule molecule molecular structural structure formula metabolite metabolism biosynthesis cycle mechanism path catabolism biosynthetic biogenesis substrate precursor synthesis derivative bioprecursor