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3-phosphoadenosine-5-phosphosulfate paps phosphosulfate synthase phosphoryl transfer papss2 papss1 sulfuryl chemical adenylyl sulfate aps phosphate protein derivative coenzyme coa 3d rendering biochemistry biology molecule molecular structural structure formula skeletal geometry iupac cas einecs metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic hmdb intermediate biogenesis substrate precursor synthesis bioprecursor