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3-hydroxyisobutyric acid 3-hydroxy-2-methylpropanoic intermediate metabolism valine chiral compound enantiomer dehydrogenase sodium salt branched chain amino cerebrospinal fluid propionyl coa methylbutyric c4h8o3 methylmalonic aminoisobutyric oxoisovalerate propanoic hydroxyisobutyric aciduria mtbe toxicity 3d rendering chemical biochemistry molecule molecular structural structure formula metabolite biosynthesis mechanism pathway catabolism biosynthetic hmdb biogenesis precursor synthesis derivative