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2-oleoylglycerol oleoylglycerol 2og monoacylglycerol monoolein gpr119 gip plasma level agonist ligand glucagon fatty acid palmotoyl endocannabinoid receptor deficiency system ecs cbr central nervous system brain neurotransmitter neuromodulatory stress appetite memory 3d rendering chemical biochemistry molecule molecular structural structure formula synthesis metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic