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molecular formula synthesis endogenous steroid catechol estrogen metabolite estradiol isomer angiogenesis 2-hydroxyestradiol 2-ohe2 metabolism ohe hydroxylation 2meoe hydroxyestrone estriol aibn methylation antioxidant genotoxicity 2-methoxyestradiol superoxide catecholaminergic production 3d rendering chemical biochemistry molecule structural structure production derivative hormone hormonal therapy replacement contraception catecholamine biosynthesis cycle mechanism pathway path hmdb