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2-c-methylerythritol 4-phosphate mep ispd arabidopsis phosphate pathway isoprenoid biosynthesis isopentenyl diphosphate isoprene diphosphocytidyl dxs mevalonate ipp ppt terpenoid precursor non-mevalonate dmapp 3d rendering chemical biochemistry organic biology molecule molecular structural structure formula skeletal geometry compound reaction synthesis derivative iupac metabolite metabolism cycle mechanism path catabolism biosynthetic hmdb intermediate biogenesis