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cdp-me 4-diphosphocytidyl-2-c-methylerythritol intermediate mep pathway non-mevalonate isoprenoid precursor biosynthesis ispd kinase ispe erythritol mevalonate independent mevalonic acid cytidine malaria methylerythritol phosphate escherichia coli cyclodiphosphate doxp 3d rendering chemical atom molecule molecular structural structure formula geometry compound metabolite metabolism cycle mechanism path catabolism biosynthetic hmdb biogenesis substrate synthesis derivative bioprecursor