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lichenes peltigeraceae foliose lecanoromycetes phycobiont bryophyta ascomycetes microscopic mosses lobariaceae thallus eukaryotes lobaria cyanobacteria lichens belonging mycobiont karelia fungi symbiotic spruce moss russia mushrooms forest ecology genus tree orzega bryophytes lichen bark association lichenized photobiont taiga ascomycete trunk lichenology photosynthesis substratum ecosystem biome jagel