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thallus genus lichens russia lichenized lichenes ecology fungi mosses lobaria lecanoromycetes photobiont ascomycetes orzega microscopic foliose phycobiont tree ascomycete peltigeraceae moss belonging cyanobacteria lobariaceae lichen bryophytes association symbiotic bryophyta mycobiont karelia taiga eukaryotes bark forest spruce mushrooms trunk lichenology photosynthesis substratum ecosystem biome jagel