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africa alert amphibian arid arthropod bands beetle bird botswana brindled broad burrow calls carnivora carnivorous coat communicate crevices dark digging diurnal dry extreme eyes family foreclaws grey habitat head herpestidae hierarchy home insectivorous large legs lepidopterans light long mammal material meerkat namibia network packs pattern plant pointed predator pups ranges reptile rock snout social south southern suricata suricate suricatta systems temperatures thermoregulate tunnels underground yellowish-brown