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portuguese millipede ommatoiulus moreleti black family julidae iberian peninsula herbivorous arthropods jointed legs diplopoda myriapods insect commerce decaying leaves dead plant australia natural enemies invasive pest indigenous south africa shipping trade voyages quiescent survive long dry summer habitat pupoid stage moults grey gonopods body segment drought autumn rains cool temperature winter humid litter shrub grasslands liverworts moss culture enemy predator introduced smooth cylindrical pairs brown tree garden