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africa aggressive amphibius animal aquatic artiodactyls barrel-shaped blood body canine columnar common dangerous even-toed eyes family feet grasses graze habitat herbivorous hippo hippopotamidae hippopotamus ivory lakes large legs loss mammal mangrove meat megaherbivore membranes mouth natural nature nictitating nostrils poaching river rivers semi-aquatic short solitary stocky sub-saharan sunscreen swamps sweat tail teeth territorial threatened torso tusks ungulate ungulates unpredictable webbed wide-opening