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africa agent anti-predator antibacterial asia bath berries bill bird black broad brown call cavity crest crested defence distinctive downy dust ecosystem epops eurasian europe family feather feed flight forager forest foul-smelling frog glade grasshopper grassland ground habitat heathland himalaya hole hoopoe hudhud insect liquid long meat medium-sized migrate monogamous moult nest nestling posture preen rain reptile rotting rounded sand savanna secretion seed smell soil solitary steppe sunbathe tail tapering thin tropical undulating upupa upupidae white wing wooded