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uroporphyrinogen iii isomer metabolic intermediate biosynthesis heme metabolite metabolism cycle mechanism pathway path catabolism biosynthetic biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous human metabolome database hmdb porphyrin tetrapyrrole preuroporphyrinogen hydroxymethylbilane uroporphyrinogen-iii cosynthase coproporphyrinogen porphyria molecule molecular formula skeletal structural structure cell biology biochemistry biomolecule bioactive biochemical 3d organic