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umbelliferone 7-hydroxycoumarin hydrangine skimmetine beta-umbelliferone ultraviolet antioxidant herbal phytochemical phytotherapy plant herb natural molecule molecular formula skeletal structural structure biochemistry biomolecule biochemical 3d biogenesis antimutagenic sunscreen umbelliferae apiaceae carrot coriander garden angelica hieracium pilosella asteraceae bigleaf hydrangea hydrangea macrophylla hydrangeaceae asafoetida giant fennel ferula communis justicia pectoralis acanthaceae 7-hydroxy-2h-1-benzopyran-2-one phenylpropanoid ultraviolet fluorescence sunscreen agent optical brightener gain medium dye lasers fluorescence indicator