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k-strophanthidin strophanthidin cardenolides adonis vernalis ranunculaceae aglycone cymarigenen ouabain membrane protein atpase muscle tissue heart diastolic dysfunction arrythmia arrow poison steroid glycoside heart-arresting cardiac inotropic effect myocardium herbal phytochemical phytotherapy plant herb natural herbalism pharmacognosy pharmacopoeia venom toxicant toxic toxicity toxin toxicology poisoning danger kill antidote molecule molecular formula skeletal structural structure 3d