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sphingosine amino alcohol unsaturated cell membrane lipids sphingolipid aliphatic phosphatidylcholine pc lecithin sphingomyelin phosphatidylethanolamine phospholipid membrane choline cell arachidonic acid lipid phosphocholine fatty ceramide lysophosphatidylcholine bilayer glycerophospholipid phosphatidylserine lipophilic signal transduction recognition neural tissue cerebrosides globosides glycosphingolipids neurotransmitter receptor neuromodulatory nervous system brain neuroscience 3d biochemistry biomolecule bioactive biochemical molecule