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somatoliberin somatocrinin growth hormone–releasing hormone ghrh gh-rh factor ghrf grf somatotropin-releasing srh factor srf somatorelin peptide arcuate nucleus hypothalamus neurosecretory neuron hypothalamo-hypophyseal portal system anterior pituitary gland hormone-releasing hormone receptor slow-wave sleep signal transduction camp-dependent pathway phospholipase somatostatin growth-hormone-inhibiting circulation 3d neurotransmitter agonist neuromodulatory nervous brain endocannabinoid stress appetite memory signaling neuroscience molecule molecular formula skeletal structural structure