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secretin water homeostasis hormone stomach pancreas liver secretions hormonal function action endocrine endocrinology level prohormone peptide cells intestinal glands duodenum sct gene prosecretin precursor gastric acid parietal cholangiocytes bile acids cholecystokinin gallbladder osmoregulation hypothalamus pituitary gland kidney inhibitory peptide small intestine regulation gastrin electrolyte water pancreatic duct epididymis molecule molecular formula skeletal structural structure 3d