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gastrin gastrin protein hormone intestine ii gastric acid hcl parietal cells stomach pyloric antrum duodenum pancreas cholecystokinin receptor peptides pepsinogen chief pancreatic gallbladder lower esophageal sphincter les gastrocolic reflex grp gastrin-releasing peptide cholinergic agent beta-adrenergic enterogastric inhibitory gip secretin somatostatin glucagon calcitonin rendering hormonal therapy replacement function action endocrine endocrinology level prohormone analogue