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pyrroloquinoline quinone pqq redox cofactor antioxidant kiwifruit breast milk quinoprotein quinoprotein glucose dehydrogenase sensor lactate mitochondrial biogenesis nicotinamide biosynthesis vitamin b14 coprinopsis cinerea heme pyridine ring metabolite metabolism cycle mechanism pathway path catabolism biosynthetic intermediate substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor cell biology biochemistry biomolecule bioactive molecule molecular formula skeletal structural 3d