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platelet-activating factor paf-acether paf agepc acetyl-glyceryl-ether-phosphorylcholine phospholipid activator mediator leukocyte platelet aggregation inflammation anaphylaxis oxidative burst chemotaxis arachidonic acid metabolism phagocytes cell endothelial cells neutrophils monocytes macrophages fungal penicillium antifungal platelet blood vessels allergic responses shock basophils polymorphonuclear lipid remodeling thrombin plasmalogen rendering biosynthesis mechanism biosynthetic intermediate biogenesis synthesis derivative bioprecursor metabolic endogenous