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phosphocreatine creatine phosphate cp pcr phosphorylated creatine skeletal muscle myocardium brain adenosine triphosphate energy enzyme kidney phosphorylcreatine creatine-p phosphagen fosfocreatine guanidinoacetate kinase adp atp diphosphate rendering metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous contractor molecule molecular formula skeletal