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phosphocholine intermediate phosphatidylcholine synthesis choline kinase atp choline adp lecithin nematodes placentas posttranslational modification c-reactive protein crp phagocytotic egg biomimetic 3d metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic biogenesis derivative endogenous contractor pc sphingomyelin phosphatidylethanolamine phospholipid membrane cholesterol fatty ceramide lysophosphatidylcholine bilayer surfactant glycerophospholipid phosphatidylserine lipophilic