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phosphatidylethanolamine pe phospholipid biological membrane phosphatidylcholine pc lecithin sphingomyelin choline cell arachidonic acid lipid phosphocholine cholesterol fatty ceramide lysophosphatidylcholine bilayer surfactant glycerophospholipid phosphatidylserine lipophilic membrane fusion curvature lipoprotein low-density lipoproteins golgi apparatus metabolite metabolism biosynthesis mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous 3d