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pancreatic polypeptide cells endocrine pancreas pancreatic liver glycogen tumour precursor icosapeptide oligopeptide neuropeptide head secretion somatostatin digestive enzyme insulin glucagon verner-morrison syndrome gastrinomas vagotomy atropine biological polymer oligomer protein macromolecule peptide peptide hormone protein 3d metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate synthesis derivative bioprecursor metabolic endogenous