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noladin ether 2-arachidonyl glyceryl 2-age endocannabinoid noladin arachidonic acid porcine brain neurotransmitter receptor agonist neuromodulatory nervous system stress appetite memory signaling neuroscience blood pressure hypotension cannabinoid metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor molecule molecular formula skeletal structural structure 3d