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met-enkephalin enkephalin opioid growth factor ogf naturally occurring endogenous peptide opioid leu-enkephalin receptor opioids pentapeptide adrenal medulla central nervous system cns striatum cerebral cortex olfactory tubercle hippocampus septum thalamus periaqueductal gray dorsal horn spinal cord biosynthesis neurotransmitter agonist neuromodulatory nervous brain signaling neuroscience metabolite metabolism cycle mechanism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous 3d