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malic acid organic compound molecular formula hydroxybutanedioic 2-hydroxysuccinic stereoisomeric food additive citric acid cycle intermediate taxonomic biochemistry calvin anaplerotic reactions phosphoenolpyruvate apple juice apricots blackberries blueberries cherries grapes mirabelles peaches pears plums quince malolactic fermentation vinegar flavors maleic anhydride enantiomer racemic plant defense rendering metabolite metabolism biosynthesis mechanism pathway path biosynthetic biogenesis precursor synthesis derivative bioprecursor metabolic enzyme