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lysophosphatidylethanolamine lpe phosphatidylethanolamine cell membranes fatty acid phospholipid egg yolk lecithin soybean signaling rendering metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme phosphatidylcholine pc sphingomyelin membrane choline arachidonic lipid phosphocholine cholesterol fatty ceramide lysophosphatidylcholine bilayer surfactant glycerophospholipid phosphatidylserine lipophilic