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lysophosphatidylcholine lpc lysopc lysolecithins phosphatidylcholine fatty acid enzyme phospholipase a2 apoptotic phagocytes multiple sclerosis myelin sheath myelin macrophages microglia phagocytose cell membrane lysophospholipase atherosclerosis 3d metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic pc lecithin sphingomyelin phosphatidylethanolamine phospholipid choline phosphocholine cholesterol glycerophospholipid phosphatidylserine