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luteinizing hormone-releasing hormone pituitary gland brain lh follicle-stimulating fsh estrogen progesterone testosterone lh-rh lhrh hypothalamus gnrh gonadotropin-releasing releasing anterior pituitary gonadotropin gnrh1 grh hh12 lnrh gnrh neuron hypothalamic–pituitary–gonadal axis synthesis hypophysial neurohormone neural cell terminal preoptic area 3d hormonal therapy replacement function action endocrine endocrinology level prohormone analogue structural structure formula