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neurokinin tachykinin peptide gonadotropin-releasing hormone nkb pregnancy kisspeptin pre-eclampsia arcuate nucleus arc neurotransmitter receptor agonist neuromodulatory nervous system brain endocannabinoid stress appetite memory signaling neuroscience metabolite metabolism biosynthesis mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous ovarian cycle gnrh luteinizing molecule molecular formula skeletal structural structure 3d