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guanine 2-aminohypoxanthine 2-amino-6-hydroxypurine nucleobase nucleic acids dna rna nucleoside guanosine adenine cytosine thymine uracil purine pyrimidine-imidazole bicyclic molecule tautomeric hydrogen bonds polymerization metabolite metabolism biosynthesis mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic endogenous guanosine phosphorylase inosine monophosphate dehydrogenase deep-sea fish reptiles crocodiles iridescent nail polish molecular formula skeletal structural structure 3d