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glutaric acid n-pyrotartaric pentanedioic propanedicarboxylic propane dicarboxylic biochemistry amino acids aciduria encephalopathy plasticizer precursor polyesters hydrogenation polyols polyamides pyrogallol safety irritation substance matter material compound component agent synthesis reaction production metabolite metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate derivative bioprecursor metabolic enzyme endogenous 3d