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formiminoglutamic acid figlu conjugate base formiminoglutamate catabolism l-histidine l-glutamic biomarker intracellular folate vitamin b12 deficiency folate liver failure disease methylmalonic homocysteine n-aminomethylidene-l-glutamic 3d metabolite metabolism biosynthesis cycle mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous contractor human metabolome database hmdb bioactive cell biochemistry biology biomolecule molecule structure formula ball stick