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etiocholanolone glucuronide etio-g endogenous metabolite testosterone liver etiocholanolone udp-glucuronyltransferases androsterone 3d hormone hormonal function endocrine endocrinology level prohormone metabolism biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme contractor bioactive biochemistry biology biomolecule molecule structural structure formula skeletal geometry ball stick