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estriol sulfate glucuronide endogenous naturally occurring diconjugated metabolite estriol liver sulfate udp-glucuronyltransferase urine pregnancy rendering mother gynecology obstetrics woman motherhood birth fertility reproduction reproductive female sex hormone menopausal hormone therapy gynecological conditions hormonal function action endocrine endocrinology level prohormone metabolism biosynthesis mechanism catabolism biosynthetic intermediate biogenesis synthesis derivative bioprecursor metabolic