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dihydrocortisol 5a-dihydrocortisol hydrallostane allodihydrocortisol metabolite cortisol 5a-reductase aqueous humor eye lens hormone hormonal therapy replacement function action endocrine endocrinology level prohormone analogue catecholamine norsteroid mineralocorticoid corticosteroid glucocorticosteroid steroid glucocorticoid receptor inflammation immune system autoimmune disease stress response carbohydrate metabolism protein catabolism anti-inflammatory molecule molecular formula skeletal structural structure geometry ball stick biomolecule bioactive 3d