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dehydroepiandrosterone sulfate dhea dhea-s androstenolone endogenous androstane steroid adrenal cortex dehydroepiandrosterone dhea neurosteroid neurotrophin nmda receptor negative allosteric modulator gaba glycine agonist bdnf brain-derived neurotrophic factor androgenic estrogenic prohormone zona reticularis adrenocorticotropic hormone acth cholesterol steroid sulfatase sts premenopausal postmenopausal prostate cancer insufficiency hypopituitarism rendering hormonal therapy replacement function action endocrine level analogue norsteroid mineralocorticoid corticosteroid steroidal