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cortisol steroid hormone hormonal therapy replacement function action endocrine endocrinology level prohormone analogue mineralocorticoid corticosteroid glucocorticosteroid glucocorticoid receptor inflammation immune system autoimmune disease stress response carbohydrate metabolism protein catabolism anti-inflammatory hydrocortisone zona fasciculata adrenal cortex gland diurnal cycle blood-glucose concentration blood sugar gluconeogenesis bone formation glucose glycogenolysis catecholamines electrolyte balance 3d molecule molecular formula skeletal structural structure