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corticoliberin corticotropin-releasing hormone crh factor crf corticotropin corticotrophin peptide stress response releasing acth hpa axis paraventricular nucleus pvn stress hypothalamus alzheimer's disease major depression corticotrope cortisol glucocorticoids mineralocorticoid adrenocorticotropic rendering pathway hormonal contractor biogenesis metabolite therapy action hmdb replacement norsteroid biosynthesis path catecholamine enzyme metabolism precursor biosynthetic cycle metabolic catabolism analogue intermediate