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coenzyme q10 ubiquinone coenzyme ubidecarenone coq10 coq vitamin ubiquinone-10 eukaryotic mitochondria electron transport chain aerobic cellular respiration redox semiquinone ubisemiquinone ubiquinol reduced oxidized iron–sulfur clusters free-radical–scavenging antioxidant radical deficiency supplement biological activity bioactive nutrient food additive nutraceutica dietary cancer dental metabolite metabolism biosynthesis cycle mechanism pathway path biosynthetic intermediate biogenesis substrate precursor synthesis metabolic enzyme 3d