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choline essential nutrient cation bilineurine vitamin b4 phospholipid phosphatidylcholine amino acid–like metabolism cell membrane organelle low-density lipoproteins acetylcholine neurotransmitter s-adenosylmethionine sam methyl donor homocysteine non-alcoholic fatty liver disease low blood pressure sweating diarrhea fish-like body odor metabolite biosynthesis cycle mechanism pathway path catabolism biosynthetic intermediate biogenesis substrate precursor synthesis derivative bioprecursor metabolic enzyme endogenous quaternary ammonium absorption glycerophosphocholine osmoregulator 3d